Physiology of hearing (ENT)
Broadly hearing mechanism can be divided into:
- Mechanical conduction of sound (done by middle ear).
- Transduction of mechanical energy into electrical impulses (done by sensory system of cochlea)
- Conduction of electrical impulse to brain (i.e. auditory pathway)
i. Conduction of sound:
- It is done mainly by middle ear. Middle ear not just simply conducts the sound but converts sound of great amplitude and less force to that of less amplitude and greater force.
- This function of the middle ear is called as impedance matching mechanism or the transformer action.
ii. Transduction of mechanical energy to electrical impulse:
- Movements of the stapes footplate causes vibrations in scala vestibuli followed by scala tympani and is transmitted to the cochlear fluids which brings about movement of the basilar membrane.
- This sets up shearing force between the tectorial membrane and the hair cells.
- The distortion of hair cells gives rise to electrical nerve impulse.
- A sound wave, depending on its frequency, reaches maximum amplitude on a particular place on the basilar membrane, and stimulates that segment (traveling wave theory of von Bekesy).
- Higher frequencies are represented in the basal turn of cochlea and the progressively lower one toward the apex.
PHYSIOLOGY OF HEARING
Conduction of sound
- Impedance matching mechanism or Transformer action of middle ear: conversion of sound of greater amplitude with lesser force to that of lesser amplitude but greater force
- It is accomplished by
- Lever action of ossicles
- Handle of malleus is 1.3 times longer than the long process of incur
- Mechanical advantage = 1.3
- Lever ratio = 1.3 : 1
Hydraulic action of TM
- Effective vibratory area of TM = 55 sq.mm.
- Area of Foot Plate of stapes = 3.2 sq.mm.
- Hydraulic ratio = 17: 1
- The product of Lever ratio and Hydraulic ratio (17 x 1.3) gives the Transformer ratio of 22: 1
Natural resonance and efficiency of auditory apparatus
- External auditory canal — 3000 Hz
- Tympanic membrane — 800 to 1600 Hz
- Middle ear — 800 Hz
- Ossicular chain — 500 to 2000 Hz
Intensity of various sound at a distance of 1 m
- Whisper — 30 dB
- Normal conversation — 60 dB
- Shout — 90 dB
- Discomfort of the ear — 120 dB
- Pain in the ear — 130 dB
Frequency range in normal hearing — 20 Hz to 20000 Hz
In routine audiometric testing — 125 to 8000 Hz are evaluated
Exam Question
- Function of Stapedius is Protects the ear from loud frequency sound.
- Exposure to noise above 160 dB causes permanent loss of hearing.
- Permanent privation of the hearing of either ear ,even though correctable by surgery,comes under Greivous injury.
- Psycho-acoustic index of loudness is indicated by the term ‘Phon’.
- Sense gravity and Linear acceleration is associated with ‘otolith organs’.
- The limit of loudness expressed as decibels that people can tolerate without substantial damage to their hearing is 85 dB.
- Foetus starts hearing by what time in intrauterine life by 20 weeks.
- Sense organ for hearing is Organ of Corti.
- Bones of middle ear are responsible for Reduction of impedance to sound transmission.
- Impedance matching occurs at Difference of surface are of tympanic membrane and foot plate.
- Primary receptor cells of hearing is Hair cell
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